Standard methods for varroa research.

Vincent Dietemann1,2*, Francesco Nazzi3, Stephen J Martin4, Denis L Anderson5, Barbara Locke6, Keith S Delaplane7, Quentin Wauquiez1, Cindy Tannahill9, Eva Frey8; Bettina Ziegelmann8, Peter Rosenkranz8, James D Ellis9.

1Swiss Bee Research Centre, Agroscope Liebefeld-Posieux Research Station ALP, Bern, Switzerland.

2Social Insect Research Group, Department of Zoology and Entomology, University of Pretoria, Pretoria, South Africa.

3Dipartimento di Scienze Agrarie e Ambientali, Università degli Studi di Udine, Italy.

4School of Environment and Life Sciences, University of Salford, Manchester, UK.  

5CSIRO Entomology, Canberra, ACT 2601, Australia.

6Department of Ecology, Swedish University of Agricultural Sciences, Uppsala, Sweden.

7Department of Entomology, University of Georgia, Athens, GA 30602, USA.

8University of Hohenheim, Apicultural State Institute, 70593 Stuttgart, Germany.

9Honey Bee Research and Extension Laboratory, Department of Entomology and Nematology, University of Florida, Gainesville, Florida, USA.

Received 16 May 2012, accepted subject to revision 25 October 2012, accepted for publication 14 November 2012.

*Corresponding author: Email:


We are grateful to Diana Sammataro and Marla Spivak for their comments on an earlier version of this paper.